UNIVERSITY PARK, Pa. — No. 18 Iowa led No. 17 Penn State, 12-0, with 5:22 left in the first quarter.

From there, the Nittany Lions outscored the Hawkeyes, 30-12, handing Iowa its second loss of the season.

Quarterback Nate Stanley was nowhere near the same passer he was against Minnesota or Indiana; the junior finished just 18-of-49 for 205 yards and 2 interceptions.

Iowa’s offense was stagnant following its shutout win over Maryland on Oct. 20, and it came back to bite the Hawkeyes. The Hawkeyes ran for 134 yards, but without any whisper of a consistent passing game, Iowa couldn’t move the ball with ease.

Mekhi Sargent was Iowa’s lone offensive spark for most of the game. He carried the ball 16 times for 91 yards (5.7 yards per carry) and did most of his damage once starter Ivory Kelly-Martin hurt his ankle and played sparingly.

Penn State quarterback Trace McSorley hurt his right knee during the second quarter, but he came back and carved up the Hawkeye defense in the second half. On Penn State’s first possession of the second half, the senior ripped off a 51-yard touchdown run. In the passing department, he finished 11-of-25 for 167 yards and 2 touchdowns.

Iowa tallied a pair of safeties against Penn State in the first half. The first came when Hawkeye Dominique Dafney blocked a punt, and the second came after Penn State snapped the ball over its punter and out of the end zone.

Iowa’s lone highlight of the game came facing a fourth-and-goal. The Hawkeyes lined up for a field goal, but holder Colten Rastetter ended up throwing the ball to defensive lineman Sam Brinks for a touchdown.

Aside from that, however, the Iowa offense did not find the end zone and had trouble capitalizing on key plays. That, ultimately, came back to bite the Hawkeyes.

Iowa had two chances to go ahead late in the fourth, but both came up zero. Stanley threw his second interception of the game on a first-and-goal inside the Penn State 5-yard line, and then he fumbled on the game’s final play.

Now, Iowa turns to Purdue. The Hawkeyes and Boilermakers will square off on Nov. 3 in West Lafayette, Indiana.